Dr Daniel Hending
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My Published Work
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In Press

2026

Hending, D. (2026) Recognizing fieldwork: Towards equitable credit in life science research. Biodiversity and Conservation. 35, 19.   Available online (Open Access)

Beutelmann, S., Rehling, F. and Hending, D. (2026) Habitat suitability modelling and environmental drivers of Malagasy fruit bats (Pteropodidae). Biodiversity and Conservation. 35, 1.    Available online (Open Access)

2025

Hending, D. (2025) Cryptic species conservation: A review. Biological Reviews. 100(1), pp. 258-274.   Available online (Open Access)

van Elst, T., Sgarlata, GM., Schüßler, D., Tiley, GP., Poelstra, JW., Scheumann, M., Blanco, MB., Aleixo-Pais, IG., Evasoa, MR., Ganzhorn, JU., Goodman, SM., Hasiniaina, AF., Hending, D., Hohenlohe, PA., Ibouroi, MT., Iribar, A., Jan, F., Kappeler, PM., Le Pors, B., Manzi, S., Olivieri, G., Rakotonanahary, AN., Rakotondranary, SJ., Rakotondravony, R., Ralison, JM., Ranaivoarisoa, JF., Randrianambinina, B., Rasoloarison, RM., Rasoloharijaona, S., Rasolondraibe, E., Teixeira, H., Zaonarivelo, JR., Louis, EE., Yoder, AD., Chikhi, L., Radespiel, U. and Salmona, S. (2025) Integrative taxonomy clarifies the evolution of a cryptic primate clade. Nature Ecology and Evolution. 9, pp. 57-72.   Available online (Open Access)

Hending, D., Randrianarison, H., Andriamavosoloarisoa, NNM., Ranohatra-Hending, C., McCabe, G., Cotton, S. and Holderied, M. (2025) Impact of forest fragmentation and associated edge-effects on tropical forest biodiversity in North West Madagascar, assessed via ecoacoustics. Animal Conservation.   Available online (Open Access)

Darras, KFA., Rountree, R., Van Wilgenburg, S., Cord, AF., Chen, Y., … Hending, D., … Xu, J., Zhou, W., Li, S., Sousa-Lima, R. and Wanger, TC. (2025) Worldwide Soundscapes: A synthesis of passive acoustic monitoring across realms. Global Ecology and Biogeography. 34(5), e70021.   Available online (Open Access)

Mulder, T., Yang, Y., Morley, E., Miller, TE., Hending, D., Taylor, GK. and Mortimer, B. (2025) Constraints on percussive seismic signals in a noisy environment by European fiddler crabs, Afruca tangeri. Journal of Experimental Biology. 228(7), jeb249323.   Available online (Open Access)

Wilmé, L., Waeber, PO., De Grave, A., Ganzhorn, JU., Llopis, JC., Kull, CA., Gardner, CJ., Salmona, J., Hending, D., Moutou, F., Razafindratsima, OJ., Radespiel, U., Lourenco, WR., Schuurman, D., Ratsimbazafy, J., Camara, C., Jenkins, PD., Ferguson, B., Raherilalao, MJ., Richard, AF., Sparks, JS. and Lowry II, PP. (2025) Position statement on the use of AI in scientific writing and publishing. Madagascar Conservation and Development. 20(1), pp. 6-9.   Available online (Open Access)

2024

Hending, D., Randrianarison, H., Andriamavosoloarisoa, NNM., Ranohatra-Hending, C., McCabe, G., Cotton, S. and Holderied, M. (2024) Forest Fragmentation and Edge Effects impact Body Condition, Fur Condition, and Ectoparasite Prevalence in a Nocturnal Lemur Community. Conservation Physiology. 12(1), coae042.   Available online (Open Access)

Hending, D.
, Randrianarison, H., Andriamavosoloarisoa, NNM., Ranohatra-Hending, C., McCabe, G., Cotton, S. and Holderied, M. (2024) Impact of Forest Fragmentation and Associated Edge Effects on the Population Density of four Nocturnal Lemur Species in North West Madagascar. Animal Conservation. 27(4), pp. 522-537.   Available online (Open Access)

Hending, D.
, Randrianarison, H., Andriamavosoloarisoa, NNM., Ranohatra-Hending, C., Cotton, S., Holderied, M. and McCabe, G. (2024) Effects of forest fragmentation on the dietary ecology and activity of a nocturnal lemur community in North West Madagascar. American Journal of Primatology. 86(1), e23569.   Available online (Open Access)

Ganzhorn, JU., Andriambeloson, JB., Atsalis, S., Behrendt, LM., Blanco, MB., Bollen,, A., Carrière, SM., Chikhi, L., Dammhahn, M., Donati, G., Eppley, T., Ernest, R., Giertz, P., Goodman, SM.,  Hending, D., Holst, F., Roberts, SH., Irwin, MT., Lahann, P., Louis, EE., Radespiel, U., Rakotondranahary, SJ., Ramanamanjato, JB., Ramananjato, V., Randriatafika, F., Ratovonamana, YR., Razafindratsima, OH., Salmona, J., Schwab, D. and Tsagnangara, C. (2024) Facilitated forest restoration using pioneer seed dispersers in Madagascar: The example of Microcebus spp. Land. 13(12), 1971.   Available online (Open Access)

Rowe, AK., Hending, D., Schäffler, L. and Markolf, M. (2024) Coquerel’s Giant Mouse Lemur, Mirza coquereli (A. Grandidier, 1867). In: Primates in Peril: The World’s 25 Most Endangered Primates 2023–2025 (Mittermeier, RA., Reuter, KE., Rylands, AB., Jerusalinsky, L., Schwitzer, C., Strier, KB., Ratsimbazafy, J. and Humle, T. Eds.). pp. 16-19. IUCN SSC Primate Specialist Group, International Primatological Society, Re:wild, Washington, DC.   Available online (Open Access)

Schüßler, D., Rabemananjara, NR., Radriarimanga, T., Rafamantanantsoa, SM., Randimbiharinirina, RD., Radespiel, U. and Hending, D. (2024) Phaner furcifer - The ghost lemur of northeastern Madagascar? Lemur News. 24, pp. 19-23.   Available online (Open Access)

Hending, D., Randrianarison, H. and Andriamavosoloarisoa, NNM. (2024) A case of hypomelanism in the northern giant mouse lemur (Mirza zaza). Lemur News. 24, pp. 2-4.   Available online (Open Access)

Schüßler, D., Blanco, MB., Guthrie, N., Sgarlata, G., Dammhahn, M., Ernest, R., Evasoa, M., Hasaniaina, A., Hending, D., Jan, F., Le Pors, B., Miller, A., Olivieri, G., Rakotonanahary, A., Rakotondranary, S., Rakotondravony, R., Ralantoharijaona, T., Ramananjato, V., Randrianambinina, B., Raeolinjanakolona, N., Rasoazanabary, E., Rasoloarison, R., Rasolofoson, D., Rasoloharijaona, S., Rasolondraibe, E., Roberts, S., Teixeira, H., van Elst, T., Johnson, S., Ganzhorn, J., Chikhi, L., Kappeler, P., Louis, E., Salmona, J. and Radespiel, U. (2024) Morphological variability or inter-observer bias? A methodological toolkit to improve data quality of multi-researcher datasets for the analysis of morphological variation. American Journal of Biological Anthropology. 183, pp. 60-78.    Available online

2023

Hending, D., Randrianarison, H., Andriamavosoloarisoa, NNM., Ranohatra-Hending, C., Holderied, M., McCabe, G. and Cotton, S. (2023) Forest fragmentation and its associated edge-effects reduce tree species diversity, size, and structural diversity in Madagascar’s transitional forests. Biodiversity and Conservation. 32, pp. 3329-3353.    Available online (Open Access)

Hending, D., Andrianiaina, A., Rakotomalala, Z. and Cotton, S. (2023) Conservation value of vanilla agroecosystems for vertebrate diversity in northeast Madagascar. Oryx. 57(1), pp. 118-128.    Available online (Open Access)

Hending, D., McCabe, G., Cotton, S. and Holderied, M. (2023) Conservation biogeography of the Dwarf Lemurs (Cheirogaleus) of Madagascar, investigated via ecological niche modelling. International Journal of Primatology. 44, pp. 960–983​.   Available online (Open Access)

Hending, D., Randrianarison, H., Andriamavosoloarisoa, NNM., Ranohatra-Hending, C., Solofondranohatra, JS., Tongasoa, HR., Ranarison, HT., Gehrke, V., Andrianirina, N., Holderied, M., McCabe, G. and Cotton, S. (In press) Seasonal differences in the encounter rate of the fat-tailed dwarf lemur (Cheirogaleus medius) in the transitional forests of North West Madagascar: implications for reliable population density assessment. International Journal of Primatology.  44, pp. 482–498.  ​Available online (Open Access)

Mulder, T., Kiuna, S., Oduor, S., Murithi, I., Markham, A. and Hending, D. (2023) Field studies: list local contributors as authors. Nature. 618, pp. 675.   Available online

Schüßler, D., Rabemananjara, NR., Radriarimanga, T., Rafamantanantsoa, SM., Randimbiharinirina, RD., Radespiel, U. and Hending, D. (2023) Habitat and ecological niche characteristics of the elusive Hairy-eared Dwarf Lemur (Allocebus trichotis) with updated occurrence and geographic range data. American Journal of Primatology. 85(4), e23473.     Available online (Open Access)

Abondano, LA., Webber, AD., Valencia, LM., Gómez-Posada, C., Hending, D., Cortes, FA. and Fuentes, N. (2023) Community-based strategies to promote primate conservation in agricultural landscapes. In: Primates in Anthropogenic Habitats (McKinney, T., Rodrigues, M. and Waters, S. Eds.). pp. 103-120. Springer, Cham, Switzerland.

2022

Hending, D., Holderied, M., McCabe, G. and Cotton, S. (2022) Effects of future climate change on the forests of Madagascar. Ecosphere. 13(4), e4017.   Available online (Open Access)

Hending, D.
, Andrianiaina, A., Rose, M., Rowlands, A., Storm, S., Holderied, MW. and Cotton, S. (2022) Vanilla bats: Insectivorous bat diversity in the vanilla agroecosystems of northeastern Madagascar. Acta Chiropterologica. 24(2), pp. 363-378.    Available online

Hending, D., Randrianarison, H., Andriamavosoloarisoa, NNM., Ranohatra-Hending, C., McCabe, G., Cotton, S. and Holderied, M. (2022) A New Population of Mouse Lemurs (Microcebus sp.) from North-western Madagascar, with population  size and density estimates. Primate Conservation. 36, pp. 103-111.   Available online (Open Access)

Randriatahina, G., Cotton, S., Hending, D., Schwitzer, C., Tinsman, J. and Volampeno, S. (2022) Blue-Eyed Black Lemur, Eulemur flavifrons (Gray 1867). In: Primates in Peril: The World’s 25 Most Endangered Primates 2022–2023 (Mittermeier, RA., Reuter, KE., Rylands, AB., Jerusalinsky, L., Schwitzer, C., Strier, KB., Ratsimbazafy, J. and Humle, T. Eds.). pp. 16-20. IUCN SSC Primate Specialist Group, International Primatological Society, Re:wild, Washington, DC.   Available online (Open Access)

2021

Hending, D. (2021) Niche-separation and conservation biogeography of Madagascar’s fork-marked lemurs (Cheirogaleidae: Phaner): Evidence of a new cryptic species? Global Ecology and Conservation. 29, e01738.   Available online (Open Access)

Hending, D. (2021) Environmental drivers of Cheirogaleidae population density: Remarkable resilience of Madagascar’s smallest lemurs to habitat degradation. Ecology and Evolution. 11, pp. 5874-5891.   Available online (Open Access)

Hending, D., Randrianarison, H., Holderied, M., McCabe, G. and Cotton, S. (2021) The kapok tree (Ceiba pentandra (L.) Gaertn, Malvaceae) as a food source for native vertebrate species during times of resource scarcity and its potential for reforestation in Madagascar. Austral Ecology. 46(8), pp. 1440-1444.   Available online (Open Access)

Hending, D., Drew, H. and Holderied, M. (2021) Habitat use of constant frequency echolocating bats in north west Madagascar with acoustic evidence for a possible new species. Acta Chiropterologica. 23(1), pp. 153-164.   Available online

Hending, D., Randrianarison, H., Cotton, S., Holderied, M. and McCabe, G. (2021) Observations of daytime activity in the nocturnal northern giant mouse lemur (Mirza zaza). Behaviour. 58, pp. 69-76.   Available online (Open Access)

2020

Hending, D., Sgarlata, GM., LePors, B., Rasolondraibe, E., Jan, F., Rakotonanahary, AN., Ralantoharijaona, TN., Debulois, S., Andrianiaina, A., Cotton, S., Rasoloharijaona, S., Zaonarivelo, JR., Andriaholinirina, NV., Chikhi, L. and Salmona, J. (2020) Distribution and conservation status of the Endangered Montagne d’Ambre fork-marked lemur (Phaner electromontis). Journal of Mammalogy. 101(4), pp. 1049-1060.   Available online

Hending, D., Seiler, M. and Stanger-Hall, KF. (2020) The vocal repertoire of the northern giant mouse lemur (Mirza zaza) in captivity. International Journal of Primatology. 41, pp. 732-763.   Available online

Hending, D., Andrianiaina, A., Maxfield, P., Rakotomalala, Z. and Cotton, S. (2020) Floral species richness, structural diversity and conservation value of vanilla agroecosystems in Madagascar. African Journal of Ecology. 58, pp. 100-111.   Available online

Emmett, C., Hending, D., Davis, A., McCabe, G. and Bray, TC. (2020) Hearing ghosts: Acoustic survey protocol and vocalization characterization for lemur leaf frogs. Herpetological Review. 51(1), pp. 24-29.

2019

Seiler, M., Hending, D. and Stanger-Hall, KF. (2019) Spacing and mate attraction: Sex-specific function of the advertisement calls in the northern giant mouse lemur (Mirza zaza). Folia Primatologica. 90, pp. 361-378.   Available online

2018

Hending, D., Andrianiaina, A., Rakotomalala, Z. and Cotton, S. (2018) The use of vanilla plantations by lemurs: encouraging findings for both lemur conservation and sustainable agroforestry in the SAVA region, north east Madagascar. International Journal of Primatology. 39(1), pp. 141-153.   Available online

Hending, D.
, McCabe, G. and Holderied, M. (2018) Mouse lemurs with “eyes bigger than their belly”: a case of primates being out-muscled by hawk-moths. Lemur News. 21, pp. 5-6.   Available online (Open Access)

2017

Hending, D., McCabe, G. and Holderied, M. (2017) Sleeping and ranging behavior of the Sambirano mouse lemur, Microcebus sambiranensis. International Journal of Primatology. 38(6), pp. 1072-1089.   Available online (Open Access)

Hending, D., Andrianiaina, A., Rakotomalala, Z. and Cotton, S. (2017) Range extension and behavioural observations of the recently described Sheth’s dwarf lemur (Cheirogaleus shethi). Folia Primatologica. 88, pp. 401-408.   Available online
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Hending, D., Holderied, M. and McCabe, G. (2017) The use of vocalizations of the Sambirano mouse lemur (Microcebus sambiranensis) in an acoustic survey of habitat preference. International Journal of Primatology. 38(4) pp. 732-750.   Available online (Open Access)

Theses and Other Selected Reports
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Hending, D., Randrianarison, H., Andriamavosoloarisoa, NNM., Ranohatra-Hending, C., Holderied, MW., McCabe, G. and Cotton, S. (2022) Spatial, behavioural and physiological responses of nocturnal lemurs (Cheirogaleidae and Lepilemuridae) to habitat fragmentation in north western Madagascar. Unpublished Report.

Hending, D. (2021) Conservation Biogeography of the Cheirogaleidae. PhD Thesis, University of Bristol.   Available online (Open Access)

Sgarlata, GM., Le Pors, B., Salmona, J., Hending, D., Chikhi, L. and Cotton, S. (2020) Phaner electromontis. The IUCN Red List of Threatened Species 2020: e.T16875A115571235.   Available online (Open Access)

Sgarlata, GM., Le Pors, B., Blanco, M., Salmona, J., Chikhi, L., Louis, EE., Cotton, S., Hending, D., Frasier, CL., Dolch, R., Ganzhorn, J., Greene, LK., Lewis, R., Rafalinirina, HA., Raharivololona, B., Rakotoarisoa, G., Ralison, J., Randriahaingo, HNT., Rasoloarison, RM., Razafindrasolo, M., Wright, P. and Zaonarivelo, J. (2020) Cheirogaleus shethi. The IUCN Red List of ThreatenedSpecies 2020: e.T163020756A163020759.   Available online (Open Access)

Randriatahina, G., Mandl, I., Rabemananjara, N., Seiler, M., Hending, D., Schwitzer, C. and Volampeno, S. (2020) Lepilemur sahamalaza. The IUCN Red List of Threatened Species 2020:e.T136645A115584072.   Available online (Open Access)

Reuter, KE., Eppley, TM., Hending, D., Pacifici, M., Semel, B. and Zaonarivelo, J. (2020) Eulemur coronatus. The IUCN Red List of Threatened Species 2020: e.T8199A115561046.    Available online (Open Access)

Sgarlata, GM., Le Pors, B., Blanco, M., Salmona, J., Chikhi, L., Louis, EE., Cotton, S., Hending, D., Dolch, R., Ganzhorn, J., Greene, LK., Lewis, R., Rafalinirina, HA., Raharivololona, B., Rakotoarisoa, G., Ralison, J., Randriahaingo, HNT., Rasoloarison, RM., Razafindrasolo, M., Wright, P. and Zaonarivelo, J. (2020) Microcebus tavaratra. The IUCN Red List of ThreatenedSpecies 2020: e.T41571A115579176.   Available online (Open Access)

Cotton, S., Hending, D. and Andrianiaina, A. (2018) Biodiversity assessment of vanilla plantations and surrounding areas in the SAVA region of Madagascar. Conference Paper: 5th European Congress of Conservation Biology.

Hending, D. (2016) Behavioural ecology of the Sambirano mouse lemur, Microcebus sambiranensis, and its implications for conservation. MSc Thesis, University of Bristol.
Ⓒ Daniel Hending
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